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Tardigrade Taxonomy &
Systematics
Below is the current systematics of the Phylum Tardigrada down to the
genus level. The systematics is compatible with the
Species Checklist by
Degma,
Guidetti & Bertolani (2009+). The
Checklist is updated several times a year by
Peter
Degma and thus
comprises an up to date list of all described valid tardigrade species.
If you want to use this taxonomy, please cite the
Checklist as requested in its Introduction.
The site was last updated on the
05-06-2022.
Phylum
Tardigrada (Spallanzani,
1777)
1.
Class Heterotardigrada
Marcus, 1927
1. Order Arthrotardigrada
Marcus, 1927
1. Family
Archechiniscidae Binda, 1978
1.
Genus Archechiniscus
Schulz, 1953
2. Family
Batillipedidae Ramazzotti,
1962
1.
Genus Batillipes
Schulz, 1953
3. Family
Coronarctidae Renaud-Mornant,
1974
1.
Genus Coronarctus
Renaud-Mornant, 1974
2. Genus
Trogloarctus Villora-Moreno,
1996
4. Family
Halechiniscidae Thulin, 1928
1. Subfamily
Dipodarctinae Pollock, 1995
1. Genus
Dipodarctus Pollock,
1995
2. Subfamily
Euclavarctinae Renaud-Mornant,
1983
1. Genus
Clavarctus Renaud-Mornant,
1983
2. Genus
Euclavarctus
Renaud-Mornant, 1975
3. Genus
Exoclavarctus
Renaud-Mornant, 1983
4. Genus
Mobjergarctus Bussau,
1982
5. Genus
Parmursa Renaud-Mornant,
1984
6. Genus
Proclavarctus
Renaud-Mornant, 1983
3. Subfamily
Florarctinae Renaud-Mornant,
1983
1. Genus
Florarctus Delamare
Deboutteville & Renaud-Mornant, 1965
2. Genus
Ligiarctus Renaud-Mornant,
1982
3. Genus
Higginsarctus Hansen
& Kristensen, 2021
4. Genus
Wingstrandarctus
Kristensen, 1984
4. Subfamily
Halechiniscinae (Thulin,
1928)
1. Genus
Chrysoarctus
Renaud-Mornant, 1984
2. Genus
Halechiniscus Richters,
1908
5. Subfamily Orzeliscinae (Schulz,
1963)
1. Genus
Mutaparadoxipus
Gross, Miller & Hochberg,
1952
2. Genus
Orzeliscus du Bois-Reymond
Marcus, 1952
3. Genus
Opydorscus
Renaud-Mornant, 1989
4. Genus
Paradoxipus Kristensen &
Higgins, 1989
6. Subfamily Styraconyxinae
Kristensen &
Renaud-Mornant, 1983
1. Genus
Angursa Pollock, 1979
2. Genus
Bathyechiniscus
Steiner, 1926
3. Genus
Cyaegharctus
Fujimoto & Jimi, 2020
4. Genus
Lepoarctus
Kristensen &
Renaud-Mornant, 1983
5. Genus
Pleocola Cantacuzène,
1951
6. Genus
Raiarctus
Renaud-Mornant, 1981
7. Genus
Rhomboarctus
Renaud-Mornant, 1984
8. Genus
Styraconyx Thulin, 1942
9. Genus
Tetrakentron Cuénot, 1892
10. Genus
Tholoarctus
Kristensen &
Renaud-Mornant, 1983
7. Subfamily Tanarctinae
Renaud-Mornant, 1980
1. Genus
Actinarctus Schulz, 1935
2. Genus
Tanarctus
Renaud-Debyser, 1959
3. Genus
Zioella
Renaud-Mornant, 1987
5. Family Neoarctidae
de Zio
Grimaldi, D'Addabbo Gallo & Morone De Lucia,
1992
1. Genus
Neoarctus
de Zio Grimaldi, D'Addabbo Gallo & Morone
De Lucia,
1992
6. Family Neostygarctidae
de Zio
Grimaldi, D'Addabbo Gallo & Morone De Lucia,
1987
1. Genus
Neostygarctus
de Zio Grimaldi, D'Addabbo Gallo & Morone
De Lucia,
1982
7. Family Renaudarctidae
Kristensen &
Higgins,
1984
1. Genus
Renaudarctus
Kristensen &
Higgins,
1984
8. Family Stygarctidae
Schulz,
1951
1.
Subfamily Megastygarctidinae
Bello & de Zio
Grimaldi,
1998
1. Genus
Megastygarcides
McKirdy,
Schmidt & McGinty-Bayly,
1976
2.
Subfamily Stygarctidinae
Schulz,
1951
1. Genus
Faroestygarctus
Hansen, Kristensen & Jørgensen, 2012
2. Genus
Hansenarctus
Fujimoto & Ohtsuka, 2019
3. Genus
Mesostygarctus
Renaud-Mornant, 1979
4. Genus
Parastygarctus
Renaud-Debyser, 1965
5. Genus
Prostygarctus
Rubal, Veiga, Fontoura & Sousa-Pinto, 2013
6. Genus
Pseudostygarctus
McKirdy,
Schmidt & McGinty-Bayly,
1976
7. Genus
Stygarctus
Schulz,
1951
2. Order Echiniscoidea
Richters, 1926
1. Family
Echiniscoididae
Krsitensen & Hallas, 1980
1.
Subfamily Echiniscoidinae
Møbjerg, Kristensen & Jørgensen,
2016
1.
Genus Anisonyches
Pollock, 1975
2.
Genus Echiniscoides
Plate, 1888
3.
Genus Neoechiniscoides
Møbjerg, Jørgensen & Kristensen,
2019
2.
Subfamily Isoechiniscoidinae
Møbjerg, Kristensen & Jørgensen,
2016
1.
Genus Isoechiniscoides
Møbjerg, Kristensen & Jørgensen,
2016
2. Family
Carphaniidae Binda &
Krsitensen, 1986
1.
Genus Carphania
Binda, 1978
3. Family Oreellidae
Puglia, 1959
1.
Genus Oreella
Murray, 1910
4. Family Echiniscidae
Thulin, 1928
1.
Genus Acanthechiniscus
Vecchi, Cesari, Bertolani,
Jönsson, Rebecchi & Guidetti, 2016
2.
Genus Antechiniscus
Kristensen, 1987
3.
Genus Barbaria
Michalczyk, Gısiorek, Morek &
Stec, 2019
4.
Genus Bryochoerus
Marcus, 1936
5.
Genus Bryodelphax
Thulin, 1928
6.
Genus Claxtonia
Gısiorek & Michalczyk,
2019
7.
Genus Cornechiniscus
Maucci & Ramazzotti, 1981
8.
Genus Diploechiniscus
Vicente, Fontoura, Cesari,
Rebecchi, Guidetti, Serrano & Bertolani, 2013
9.
Genus Echiniscus
Schultze, 1840
10.
Genus Hypechiniscus
Thulin, 1928
11.
Genus Kristenseniscus
Gısiorek, Morek, Stec &
Michalczyk, 2019
12.
Genus Mopsechiniscus
du Bois-Reymond Marcus,
1944
13.
Genus Multipseudechiniscus
Schulte & Miller, 2011
14.
Genus Nebularmis
Gısiorek & Michalczyk,
2019
15.
Genus Novechiniscus
Kristensen, 1987
16.
Genus Parechiniscus
Cuénot,
1926
17.
Genus Proechiniscus
Kristensen, 1987
18.
Genus Pseudechiniscus
Thulin, 1911
1.
Subgenus Meridioniscus
Gısiorek,
Vonèina & Michalczyk,
2021
2.
Subgenus Pseudechiniscus
Thulin, 1911
19.
Genus Stellariscus
Gısiorek, Suzuki, Kristensen & Michalczyk, 2018
20.
Genus Testechiniscus
Kristensen, 1987
21.
Genus Viridiscus
Gısiorek & Michalczyk,
2019
22.
Genus Zealandiscus
Kaczmarek & Roszkowska,
2021
2.
Class Mesotardigrada
Rahm, 1937 (nomen dubium)
1. Order Thermozodia
Ramazzotti & Maucci, 1983 (nomen
dubium)
1. Family
Thermozodiidae Rahm, 1937
(nomen dubium)
1.
Genus Thermozodium
Rahm, 1937 (nomen dubium)
3.
Class Eutardigrada
Richters, 1926
1. Order Apochela
Schuster, Nelson, Grigarick &
Christenberry, 1980
1. Family
Milnesiidae Ramazzotti, 1962
1.
Genus Bergtrollus
Dastych, 2011
2. Genus
Limmenius Horning,
Schuster & Grigarick, 1978
3. Genus
Milnesioides Claxton,
1999
4. Genus
Milnesium Doyère,
1840
2. Order Parachela
Schuster, Nelson, Grigarick &
Christenberry, 1980
1. Superfamily Eohypsibioidea
Bertolani & Kristensen, 1987
1. Family
Eohypsibiidae
Bertolani & Kristensen, 1987
1.
Genus Austeruseus
Trygvadóttir & Kristensen,
2011
2. Genus
Bertolanius
Özdikmen,
2008
3. Genus
Eohypsibius
Kristensen,
1982
2. Superfamily
Hypsibioidea Pilato, 1969
1. Family
Calohypsibiidae Pilato, 1969
1.
Genus Calohypsibius
Thulin,
1928
2. Family
Hypsibiidae Pilato, 1969
1. Subfamily
Diphasconinae Dastych, 1992
1. Genus
Diphascon
Plate,
1889
2.
Subfamily Hypsibiinae Pilato,
1969
1.
Genus Borealibius
Pilato,
Guidetti, Rebecchi, Lisi, Hansen & Bertolani,
2006
2.
Genus Cryobiotus
Dastych,
2019
3.
Genus Hypsibius
Ehrenberg, 1848
3.
Subfamily Itaquasconinae Rudescu,
1964
1.
Genus Adropion
Pilato,
1987
2. Genus
Astatumen Pilato,
1997
3.
Genus Bindius
Pilato,
2009
4.
Genus Guidettion
Gısiorek & Michalczyk,
2020
5.
Genus Insulobius
Gısiorek & Michalczyk,
2020
6.
Genus Itaquascon
de Barros, 1939
7.
Genus Mesocrista
Pilato, 1987
8.
Genus Parascon
Pilato & Binda, 1987
9.
Genus Platicrista
Pilato, 1987
10.
Genus Raribius
Gısiorek & Michalczyk,
2020
11.
Genus Sarascon
Guil, Rodrigo & Machordom, 2015
4.
Subfamily Pilatobiinae
Bertolani, Guidetti, Marchioro, Altiero, Rebecchi & Cesari, 2014
1.
Genus Notahypsibius
Tumanov, 2020
2.
Genus Pilatobius
Bertolani, Guidetti, Marchioro,
Altiero, Rebecchi & Cesari, 2014
?.
Subfamily ?
1. Genus
Acutuncus Pilato & Binda,
1997
2.
Genus Mixibius
Pilato,
1992
3. Family
Microhypsibiidae Pilato,
1998
1.
Genus Microhypsibius
Thulin,
1928
4. Family
Ramazzottiidae Sands,
McInnes, Marley, Goodall-Copestake, Convey & Linse, 2008
1.
Genus Cryoconicus
Zawierucha, Stec, Lachowska-Cierlik, Takeuchi, Li & Michalczyk,
2018
2.
Genus Hebesuncus
Pilato,
1987
3.
Genus Ramazzottius
Binda & Pilato,
1986
3. Superfamily
Isohypsibioidea Sands, McInnes,
Marley, Goodall-Copestake, Convey & Linse, 2008
1. Family
Doryphoribiidae
Gısiorek, Stec, Morek &
Michalczyk, 2019
1.
Genus Apodibius
Dastych,
1983
2.
Genus Doryphoribius
Pilato,
1969
3.
Genus Grevenius
Gısiorek, Stec, Morek &
Michalczyk, 2019
4.
Genus Vladimirobius
Kaczmarek, Bartylak & Roszkowska, 2020
5.
Genus Paradiphascon
Dastych, 1992
6.
Genus Pseudobiotus
Nelson,
1980
7.
Genus Thulinius
Bertolani,
2003
2. Family
Halobiotidae
Gısiorek, Stec, Morek &
Michalczyk, 2019
1.
Genus Halobiotus
Kristensen,
1982
3. Family
Hexapodibiidae
Cesari, Vecchi, Palmer,
Bertolani, Pilato, Rebecchi & Guidetti, 2016
1. Genus
Haplohexapodibius
Pilato & Beasley,
1987
2.
Genus Haplomacrobiotus
May,
1948
3.
Genus Hexapodibius
Pilato,
1969
4.
Genus Parhexapodibius
Pilato,
1969
4. Family
Isohypsibiidae Sands, McInnes,
Marley, Goodall-Copestake, Convey & Linse, 2008
1.
Genus Dastychius
Pilato,
2013
2.
Genus Dianea
Gısiorek, Stec, Morek &
Michalczyk, 2019
3.
Genus Eremobiotus
Biserov,
1992
4.
Genus Fractonotus
Pilato,
1998
5.
Genus Isohypsibius
Thulin,
1928
6.
Genus Thalerius
Dastych,
2009
7.
Genus Ursulinius
Gısiorek, Stec, Morek &
Michalczyk, 2019
8.
Genus Weglarskobius
Kaczmarek, Bartylak & Roszkowska, 2020
5. Family
Ramajendidae Tumanov, 2021
1.
Genus Ramajendas
Pilato
& Binda,
1990
4. Superfamily
Macrobiotoidea Thulin, 1928
1. Family
Adorybiotidae Stec,
Vecchi &
Michalczyk, 2020
1.
Genus Adorybiotus
Maucci & Ramazzotti,
1981
2.
Genus Crenubiotus
Lisi, Londoño & Quiroga,
2020
2. Family
Macrobiotidae Thulin, 1928
1.
Genus Biserovus
Guidetti & Pilato,
2003
2.
Genus Calcarobiotus
Dastych,
1993
1.
Subgenus Calcarobiotus
Dastych,
1993
2.
Subgenus Discrepunguis
Guidetti & Bertolani,
2001
3.
Genus Famelobiotus
Pilato, Binda & Lisi,
2004
4.
Genus Insuetifurca
Guidetti & Pilato,
2003
5.
Genus Macrobiotus
Schultze,
1834
6.
Genus Mesobiotus
Vecchi, Cesari, Bertolani, Jönsson, Rebecchi &
Guidetti,
2016
7.
Genus Minibiotus
Schuster,
1980
8.
Genus Minilentus
Guidetti & Pilato,
2003
9.
Genus Paramacrobiotus
Guidetti, Schill,
Bertolani, Dandekar & Wolf,
2009
10.
Genus Pseudodiphascon
Ramazzotti,
1965 (genus dubius)
11.
Genus Pseudohexapodibius
Bertolani & Biserov,
1996
12.
Genus Schusterius
Kaczmarek & Michalczyk,
2006
13.
Genus Sisubiotus
Stec, Vecchi, Calhim & Michalczyk,
2021
14.
Genus Tenuibiotus
Pilato & Lisi,
2011
3. Family
Murrayidae Guidetti,
Rebecchi & Bertolani, 2000
1.
Genus Dactylobiotus
Schuster,
1980
2.
Genus Macroversum
Pilato &
Catanzaro,
1988
3.
Genus Murrayon
Bertolani & Pilato,
1988
4. Family
Richtersiusidae
Guidetti, Schill, Giovannini, Massa, Goldoni, Ebel, Förschler, Rebecchi & Cesari, 2021
1.
Genus Diaforobiotus
Guidetti, Rebecchi, Bertolani,
Jönsson, Kristensen & Cesari, 2016
2.
Genus Richtersius
Pilato & Binda,
1989
?. Superfamily
?
?. Family
Necopinatidae Ramazzotti & Maucci,
1983
1.
Genus Necopinatum
Pilato,
1971
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